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in biology, Tomarctus is a canid genus of the extinct subfamily Borophaginae which inhabited most of North America during the late Early Miocene to the Early Barstovian age of the Middle Miocene (23—16 million years ago). Tomarctus existed for approximately 6.83 million years.
A relative of such genera as Aelurodon and Borophagus, Tomarctus is another one of the hyena-like ‘bone crushing’ dogs of the Miocene. This description comes from the attachments for powerful jaw closing muscles and short muzzle which means that when borophagine canids like Tomarctus bit on something, it was closer to the point of jaw articulation. This in turn means that the force from the muscles is focused so that it is concentrated upon a smaller area so that more damage could be done to whatever is being bitten. This is exactly the same with a modern hyena.
The high bite force of Tomarctus is beyond that necessary to kill an animal, which leads to speculation that scavenging was just as an important part of the diet as was hunting, perhaps even more so. Scavengers however usually get to a carcass after most if not all of the fleshy parts have been devoured, probably by the animal or animal that made the kill. This would be no problem for an animal with jaws like those of Tomarctus however since it had the necessary jaw adaptations for cracking open the bones so that it could get at the marrow within. Bone marrow by itself is one of the most nutritious things in the natural world, and when encased in bone under the right conditions it can last for months and even years after the death of an animal. However, just because Tomarctus was well suited to being a scavenger, it does not mean that they never killed their own prey.
Aelurodon francisi
Aelurodon simulans
The genus currently contains two accepted species, Tomarctus brevirostris and Tomarctus hippophaga.
Tomarctus brevirostris, synonymous with Aelurodon francisi, was named by Edward Drinker Cope in 1873. Fossil specimens have been found as far south as Panama, east to Plum Point, Maryland, west to California, and north to Montana.
Tomarctus hippophaga was first described by Matthew and Cook in 1909 from the Trojan Quarry, Olcott Formation, Nebraska. Specimens have since been found as far west as California and as far north as the Montana/Alberta, Canada line.